Koko, the western lowland gorilla that began to learn sign language in the San Francisco Zoo and eventually became a celebrity beloved by millions, passed away in her sleep Tuesday at the age of 46.

Koko, the gorilla who mastered sign language, has died. The Gorilla Foundation says the 46-year-old western lowland gorilla died in its sleep at the foundation’s preserve in California’s Sana Cruz ...

Koko the gorilla, whose remarkable sign-language ability and motherly attachment to pet cats helped change the world’s views about the intelligence of animals and their capacity for empathy, has died ...

Koko, the gorilla who mastered sign language, died at the age of 46. Koko, the gorilla who mastered sign language and became a pop-culture phenomenon, will be laid to rest Saturday in a ceremony at an ...

Koko the gorilla is famous for having learned a number of hand signs to help her communicate with her caregivers. So when she was introduced to a box full of kittens she told her trainers, in the only ...

WOODSIDE, Calif. -- Koko the gorilla passed away Tuesday morning in her sleep at the age of 46. Koko was known for her extraordinary mastery of sign language. Koko was born at the San Francisco Zoo ...

WOODSIDE, Calif. -- Koko, the gorilla who mastered sign language, has died. The Gorilla Foundation says the 46-year-old western lowland gorilla died in her sleep at the foundation's preserve in ...

SAN FRANCISCO—Koko the gorilla, whose remarkable sign-language ability and motherly attachment to pet cats helped change the world’s views about the intelligence of animals and their capacity for ...

The Gorilla Foundation says the 46-year-old western lowland gorilla died in her sleep at the foundation’s preserve in California’s Santa Cruz mountains on Tuesday.

WOODSIDE, Calif. — Koko the gorilla who mastered sign language, raised kittens and once playfully tried on the glasses of the late actor Robin Williams, has died. She was 46. The Gorilla Foundation ...